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AMPHICOELIAS

a diplodocoid sauropod dinosaur from the Late Cretaceous of North America.
amphicoelias
Pronunciation: AM-fi-SEEL-ee-as
Meaning: Double hollow (biconcave)
Author/s: Cope (1877)
Synonyms: None known
First Discovery: Colorado, USA
Discovery Chart Position: #54

Amphicoelias altus

Amphicoelias altus, a diplodocid around 18 meters long and 18 tons in weight, was named by palaeontologist Edward Drinker Cope in December 1877 for two vertebrae, a pubis (hip bone), and a femur (upper leg bone) discovered in stratigraphic zone 6 of Colorado's Morrison Formation. But the real story lies with a much larger species...

Amphicoelias fragillimus

Amphicoelias fragillimus was shaping up to be the biggest dinosaur ever, and eclipsed even the Blue Whale in length when applying proportionate upscaling based on better-represented relatives to Edward Drinker Cope's sketches of a partial colossal vertebra found by Oramel Lucas at Colorado's Garden Park in 1877. Unfortunately, despite its humongous size, it was "misplaced" along with an equally huge partial femur found a short distance away, and all attempts to locate it have been in vain.

It's not beyond the realms of possibility that Cope, obsessed with one-upping his arch rival during their infamous machismo-driven bone wars, was telling fibs. But reputations were at stake, and the fact that Othniel Charles Marsh, who often sent spies to keep tabs on Cope and his discoveries, didn't publicly question the measurements of his find speaks volumes for its validity. Given its "fragillimus" state and a lack of fossil-hardening preservatives available at the time, it may have just crumbled to mush where it lay.

In 1921, Amphicoelias fragillimus was provisionally synonymised with Amphicoelias altus by H.F. Osborn and C. C. Mook, who noted similarities to Diplodocus. Then, in 2007, John Foster rocked palaeontology to its core when he suggested that these similarities were actually identicalities and that Diplodocus should probably be synonymised too. The cross-section of the thigh bone, which palaeontologists clung to as a means to separate Amphicoelias altus from Diplodocus, is now known to occur in Diplodocus too, and if they do represent the same critter, then the latter name would be sunk as the former was coined first.

The vertebra of Amphicoelias fragillimus, however, sports features not found in either Amphicoelias altus or Diplodocus and is no closer to one than it is to the other. Separated at the species level by Cope, the former may actually represent a different type of sauropod entirely, but before it can be granted its own name, the original remains need to be pinpointed for closer scrutiny, and the chances are slim... or so you might think.

Despite opining the difficulty in confirming its taxonomic validity without the holotype in hand for further study and the holotype remaining well and truly lost, Kenneth Carpenter renamed Amphicoelias fragillimus to Maraapunisaurus in 2018, reclassified it as a rebbachisaurid, and revised its vertebrae to a less spectacular 2.4 meters in height which, based on the proportions of its new-found family members, scythed close to 50% off its popular estimated body length of 58 metres.
Amphicoelias latus
Amphicoelias latus was found at the same site as Amphicoelias altus, whose name is virtually identical. It is possible that Cope confused himself and accidentally flip-flopped the first two letters of their epithets as he scribbled their names in his notebook, but he did assign the former its own holotype (AMNH 5765); four caudal vertebrae and a right femur which, despite being extraordinarily robust, has been misplaced. Nevertheless, the bits that were available to study are closer in form to those of Camarasaurus than to those of Amphicoelias, which led Osborn and Mook to assign them to Camarasaurus supremus in 1921. However, it appears to have more in common with Camarasaurus grandis.
"Amphicoelias brontodiplodocus"
In 2010, a monograph was made available (but not formally published) by Henry Galiano and Raimund Albersdorfer in which they referred a new species to AmphicoeliasAmphicoelias brontodiplodocus—based on several complete specimens found in the Dana Quarry of Big Horn Basin. Shockingly, this "paper" sunk every Morrison formation diplodocid plus a few from elsewhere into Amphicoelias altus, and the only other diplodocid the authors considered valid was their new and ridiculously named critter—literally "Both sides hollow thunder double beam". Unsurprisingly, this theory didn't receive much support from palaeontologists, which in hindsight was justified. In exchange for a bagful of cash, a specimen named "Sleeping Beauty" found itself mounted in a Dubai shopping centre in 2014 with the unbelievably mundane "DubaiDino" hanging around its neck after Saudi banker Johara Al Bayedh won a naming contest sponsored by the Emaar Malls Group. And guess what? It's a confirmed species of Diplodocus longus, the only mounted specimen made up entirely of original fossilised bone, no less. It's such a shame that kids will be monkey-swinging from its ribs and carving their names into its toes while it should be in a museum, undergoing serious scientific study.
(Elevated Double hollow)Etymology
Amphicoelias is derived from the Greek "amphi" (both sides), "koilos" (hollow) and "ias" (in character) in reference to the biconcave form of its vertebra.
The Species epithet, altus, means "high" or "elevated" in Latin.
The Species epithet, fragillimus, means "very fragile" in Latin.
Discovery
The fossils of both A. altus and A. fragillimus were recovered from Colorado's Morrison formation in 1877. Cope originally called this "the Dakota formation".
Holotype:
A. altus (AMHD 5764) consists of two back vertebrae, a hip bone (pubis), and a femur, found by Aaron Ripley (Oramel Lucas' brother-in-law) in Quarry 12.
A. fragmilis (AMNH 5777) is a single colossal vertebra, found by Oramel Lucas at a site now affectiontely known as "Cope's Nipple" near the main Camarasaurus supremus localities in Garden Park, north of Cañon City, Colorado. Despite having a number reserved for it by the AMNH, records show that it never arrived in New York! Cope's field notes contain an entry for an "immense distal end of femur" located only tens of meters away from the vertebra which might have belonged to the same individual... or to a large specimen of any of the area's other sauropods... but it was never collected.
Estimations
Timeline:
Era: Mesozoic
Epoch: Late Jurassic
Stage: Oxfordian-Kimmeridgian
Age range: 156-151 mya
Stats:
Est. max. length: 58 meters
Est. max. hip height: 9 meters
Est. max. weight: 120 tons
Diet: Herbivore
References
• Cope ED (Dec 10th, 1877) "On Amphicoelias, a genus of Saurians from the Dakota epoch of Colorado". Paleontological Bulletin no. 27. Page 2.
• Cope ED (Dec 21st, 1877) "On the Vertebrata of the Dakota Epoch of Colorado". Proceedings of the American Philosophical Society, 17(100): 233-247.
• Cope ED (August 1878) "A new species of Amphicoelias". The American Naturalist, 12(8).
• Cope ED (Feb 1st, 1878) "On the Saurians Recently Discovered in the Dakota Beds of Colorado". The American Naturalist. Printed Jan 31st, 1878. Page 82.
• Osborn HF and Mook CC (Jan. 1921) "Camarasaurus, Amphicoelias and other sauropods of Cope". Memoirs of the American Museum of Natural History, volume 3, part 3.
• McIntosh JS (1998) "New information about the Cope collection of sauropods from the Garden Park, Colorado". In Carpenter, Chure and Kirkland (eds.) "The Morrison Formation: An Interdisciplinary Study. Part 2". Modern Geology, 23(1–4): 481–506.
• Tidwell and Carpenter (2005) "Thunder-Lizards: The Sauropodomorph Dinosaurs".
• Carpenter K (2006) "Biggest of the Big: A Critical Re-Evaluation of the Mega-Sauropod Amphicoelias fragillimus Cope, 1878". New Mexico Museum of Natural History and Science Bulletin, 36: 131-138.
• Foster J (2007) "Jurassic West: Dinosaurs of the Morrison Formation and their world".
• Galiano H and Albersdörfer R (2010) "A new basal diplodocid species, Amphicoelias brontodiplodocus from the Morrison Formation, Big Horn Basin, Wyoming, with a taxonomic reevaluation of Diplodocus, Apatosaurus, Barosaurus and other genera".
• Woodruff C and Foster JR (2015) "The fragile legacy of Amphicoelias fragillimus (Dinosauria: Sauropoda; Morrison Formation – Latest Jurassic)". PeerJ PrePrints, 3: e838v1. DOI: 10.7287/peerj.preprints.838v1
• Carpenter K (2018) "Maraapunisaurus fragillimus, N.G. (formerly Amphicoelias fragillimus), a basal Rebbachisaurid from the Morrison Formation (Upper Jurassic) of Colorado". Geology of the Intermountain West, 5: 227–244. DOI: 10.31711/giw.v5i0.28.
• Paul GS (2019) "Determining the Largest Known Land Animal: A Critical Comparison of Differing Methods for Restoring the Volume and Mass of Extinct Animals". Annals of Carnegie Museum, 85(4): 335. DOI: 10.2992/007.085.0403.
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To cite this page:
Atkinson, L. "AMPHICOELIAS :: from DinoChecker's dinosaur archive".
›. Web access: 06th Mar 2026.
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