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UNENLAGIA

an omnivorous unenlagiine theropod dinosaur from the Late Cretaceous of Argentina.
unenlagia.png
Pronunciation: oon-en-LAHG-ee-a
Meaning: Half bird
Author/s: Novas and Puerta (1997)
Synonyms: None known
First Discovery: Rio Negro, Argentina
Discovery Chart Position: #420

Unenlagia comahuensis

Unenlagia was originally hailed as an early bird. Actually, it was hailed as half a bird — from the Mapudungun "uñen" (half) and "lag" (bird) — reflecting its liminal status somewhere between small theropods and "true" birds... which are small theropods. It sports a curious suite of features, as you would expect from a so-called "transitional" critter. But it's the position of its wings that has courted the most attention and put several groups of palaeontologists in a flap.

Initially, the shoulder girdle of Unenlagia was thought to be quite supple and capable of a broad range of movements. While it lacked the freedom required for Yoga or kung fu, the scapula (shoulder blade), positioned atop the rib cage and parallel to the spine, was capable of an up-and-down motion, which mirrors the flapping seen in modern birds. Later, some palaeontologists revised this view by lowering the shoulder joint and positioning the scapula upwards and across the ribcage, in a more orthodox non-bird theropod style. But that would mean the wings couldn't be lifted above its back and so a full flap, and thus powered flight, was unpossible.

With an estimated length of at least 3 meters, Unenlagia was probably too big to fly, regardless of its shoulder design. But it does belong to a family (Unenlagiinae) that contains a flying member in the form of Rahonavis, which suggests one of two things: (1) unenlagiines were flyers and Unenlagia represents a flightless lineage of oddballs, or (2) unenlagiines were flightless and Rahonavis were the evolutionary oddballs that took to the skies.
(Half bird from Comahue)Etymology
In the Mapudungun (Earth language) of the Mapuche (Earth people) — one of the indigenous inhabitants of south-central Chile and southwestern Argentina — Unenlagia is derived from "uñen" (half) and "lag" (bird) because the describers considered it to be a stepping stone between birds and basal theropods.
The species epithet, comahuensis (KOH-muh-hoo-EN-sis) honours the Comahue region of Argentina where the fossils were found.
Discovery
The first fossils of Unenlagia were discovered in the Portezuelo Formation, Sierra del Portezuelo, Neuquen Province, Comahue, Argentina in 1996.
The holotype (MCF PVPH 78) is a partial skeleton, lacking the skull but including vertebrae, a sacrum (fused hip vertebrae), ribs, a scapula (shoulder blade), a humerus (upper arm bone), a partial pelvis, a femur (thigh) and a tibia (shin).
Estimations
Timeline:
Era: Mesozoic
Epoch: Late Cretaceous
Stage: Turonian
Age range: 91-88 mya
Stats:
Est. max. length: 2.5 meters
Est. max. hip height: ?
Est. max. weight: 35 Kg
Diet: Carnivore
Unenlagia paynemili
Unenlagia paynemili was found at the "Futalognko" site near the Barreales lake (Lago Barreales), Loma de La Lata, in 2002 and named in honour of Maximino Paynemil, the chief of the Paynemil Indians that inhabit this area. Its bones — MUCPv-349; a left humerus (upper arm) and a hip bone (left pubis), MUCPv-416; a back vertebra and MUCPv-409; another hip bone (a right ilium) — are mostly more gracile than those of Unenlagia comahuensis. A toe bone (MUCPv-415) and claw (MUCPv-343) were also assigned here, albeit with some reservations. The "foot" claw may actually belongs to its hand.
References
• Novas FE and Puerta PF (1997) "New evidence concerning avian origins from the Late Cretaceous of Patagonia". Nature, 387: 390–392. DOI: 10.1038/387390a0.
• Calvo JO, Porfiri JD and Kellner AWA (2004) "On a new Maniraptoran dinosaur from the Late Cretaceous of Neuquén, Patagonia, Argentina". Arquivos do Museu Nacional, Rio de Janeiro, 62(4): 549-566.
• Norell MA and Makovicky PJ (2004) "Dromaeosauridae". In Weishampel, Dodson and Osmólska (eds.) "The Dinosauria: Second Edition".
• Senter P (2006) "Scapular orientation in theropods and basal birds, and the origin of flapping flight". Acta Palaeontologica Polonica, 51(2).
• Agnolin FL and Novas FE (2011) "Unenlagiid theropods: are they members of the Dromaeosauridae?" Anais da Academia Brasileira de Ciências, 83(1): 117-62. DOI: 10.1590/S0001-37652011000100008.
• Paul GS (2002) "Dinosaurs of the Air: The Evolution and Loss of Flight in Dinosaurs and Birds".
• Gianechini FA and Apesteguía S (2011) "Unenlagiinae revisited: dromaeosaurid theropods from South America". Anais da Academia Brasileira de Ciências, 83(1): 163-95. DOI: 10.1590/S0001-37652011000100009.
• Novas FE, Egli FB, Agnolin F and Motta MJ (2021) "Osteology of Unenlagia comahuensis (Theropoda, Paraves, Unenlagiidae) from the Late Cretaceous of Patagonia". The Anatomical Record: Advances in Integrative Anatomy and Evolutionary Biology, 304(1). DOI: 10.1002/ar.24641.
• Federico A. Gianechini FA and Zurriaguz V (2021) "Vertebral pneumaticity of the paravian theropod Unenlagia comahuensis, from the Upper Cretaceous of Patagonia, Argentina". Cretaceous Research, 127(4): 104925. DOI: 10.1016/j.cretres.2021.104925.
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To cite this page:
Atkinson, L. "UNENLAGIA :: from DinoChecker's dinosaur archive".
›. Web access: 06th Mar 2026.
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